We
investigate the time-reversal symmetry of nonequilibrium
fluctuations. Their dynamical randomness can be characterized in terms
of the standard and time-reversal entropies per unit time of dynamical
systems theory. In particular, we present experimental results for a
Brownian particle in a moving trap, showing that their difference
equals
the thermodynamic entropy production. This shows that the thermodynamic
entropy production finds its origin in the time asymmetry of the
dynamical randomness. Such concepts also give new insights to the
physics of information, such as Landauer's principle.